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The queen upon emergence of teh 3 development stages is a virgin queen. She gains strength and eventually performs orientation flights then leaves for her mating flight. Following mating (often in first week usually within first three weeks of adult life) she is termed the mated queen. Colonies normally have a single mated queen (but in a percentage of colonies there may be a mother-daughter queen in the same hive).
Queens are light aversive. Thus, when hives are opened, a queen may abandon normal behavior of egg laying and run on the comb, or run off brood combs to a comb with honey stores or to internal hive surfaces. Each hive opening causes some disturbance, complicated further by use of smoke that may disrupt pheromonepheromone:
a chemical substance released externally by an individual (from an exocrine gland) which stimulates a response in a second individual of the same species
distribution. Africanized queens and workers exhibit pronounced comb running behavior when their colonies are examined.
Poorly mated queen
There is no accepted definition of improperly (poorly) mated queens or of an inferior queen. Some commonly listed characteristics of improperly mated/inferior (and/or failing) queens are:
Delaplane, 2019. Biology of individual Honey Bees. Bee Health. https://bee-health.extension.org/biology-of-individual-honey-bees/?msclkid=c3cb798ccfcc11ecb5773b5bfd6f1e2d
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| EVENTNAME | [empty string] | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
| EXCEPTION |
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| Error - struct | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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| Detail | File or directory E:\inetpub\wwwroot\test_idtools_org\tools\1077\images\thumbnails\retinue_of_marked,_clipped_queen_R_Snyder_1.jpg does not exist. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
| Message | An exception occurred while trying to read the image. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
| StackTrace | coldfusion.image.core.ImageExceptions$ImageReadingException: An exception occurred while trying to read the image. at coldfusion.image.Image.getFilePath(Image.java:495) at coldfusion.image.Image.<init>(Image.java:372) at coldfusion.tagext.image.ImageTag.doStartTag(ImageTag.java:377) at coldfusion.runtime.CfJspPage._emptyTcfTag(CfJspPage.java:5274) at cffactSheet2ecfm54488476._factor56(E:\inetpub\wwwroot\test_idtools_org\tools\toolThemes\1\factSheet.cfm:1465) at cffactSheet2ecfm54488476._factor57(E:\inetpub\wwwroot\test_idtools_org\tools\toolThemes\1\factSheet.cfm:1452) at cffactSheet2ecfm54488476._factor58(E:\inetpub\wwwroot\test_idtools_org\tools\toolThemes\1\factSheet.cfm:1443) at cffactSheet2ecfm54488476._factor59(E:\inetpub\wwwroot\test_idtools_org\tools\toolThemes\1\factSheet.cfm:1438) at cffactSheet2ecfm54488476._factor62(E:\inetpub\wwwroot\test_idtools_org\tools\toolThemes\1\factSheet.cfm:1437) at cffactSheet2ecfm54488476._factor65(E:\inetpub\wwwroot\test_idtools_org\tools\toolThemes\1\factSheet.cfm:1284) at cffactSheet2ecfm54488476.runPage(E:\inetpub\wwwroot\test_idtools_org\tools\toolThemes\1\factSheet.cfm:1) at coldfusion.runtime.CfJspPage.invoke(CfJspPage.java:257) at coldfusion.tagext.lang.IncludeTag.handlePageInvoke(IncludeTag.java:749) at coldfusion.tagext.lang.IncludeTag.doStartTag(IncludeTag.java:578) at coldfusion.runtime.CfJspPage._emptyTcfTag(CfJspPage.java:5274) at cfindex2ecfm959615551._factor8(E:\inetpub\wwwroot\test_idtools_org\tools\toolThemes\1\index.cfm:194) at cfindex2ecfm959615551.runPage(E:\inetpub\wwwroot\test_idtools_org\tools\toolThemes\1\index.cfm:1) at coldfusion.runtime.CfJspPage.invoke(CfJspPage.java:257) at coldfusion.tagext.lang.IncludeTag.handlePageInvoke(IncludeTag.java:749) at coldfusion.tagext.lang.IncludeTag.doStartTag(IncludeTag.java:578) at coldfusion.runtime.CfJspPage._emptyTcfTag(CfJspPage.java:5274) at cfindex2ecfm1087141064.runPage(E:\inetpub\wwwroot\test_idtools_org\tools\1077\index.cfm:130) at coldfusion.runtime.CfJspPage.invoke(CfJspPage.java:257) at coldfusion.tagext.lang.IncludeTag.handlePageInvoke(IncludeTag.java:749) at coldfusion.tagext.lang.IncludeTag.doStartTag(IncludeTag.java:578) at coldfusion.filter.CfincludeFilter.invoke(CfincludeFilter.java:65) at coldfusion.filter.ApplicationFilter.invoke(ApplicationFilter.java:573) at coldfusion.filter.RequestMonitorFilter.invoke(RequestMonitorFilter.java:43) at coldfusion.filter.MonitoringFilter.invoke(MonitoringFilter.java:40) at coldfusion.filter.PathFilter.invoke(PathFilter.java:162) at coldfusion.filter.IpFilter.invoke(IpFilter.java:45) at coldfusion.filter.ExceptionFilter.invoke(ExceptionFilter.java:97) at coldfusion.filter.BrowserDebugFilter.invoke(BrowserDebugFilter.java:81) at coldfusion.filter.ClientScopePersistenceFilter.invoke(ClientScopePersistenceFilter.java:28) at coldfusion.filter.BrowserFilter.invoke(BrowserFilter.java:38) at coldfusion.filter.NoCacheFilter.invoke(NoCacheFilter.java:60) at coldfusion.filter.GlobalsFilter.invoke(GlobalsFilter.java:38) at coldfusion.filter.DatasourceFilter.invoke(DatasourceFilter.java:22) at coldfusion.filter.CachingFilter.invoke(CachingFilter.java:62) at coldfusion.CfmServlet.service(CfmServlet.java:231) at coldfusion.bootstrap.BootstrapServlet.service(BootstrapServlet.java:311) at org.apache.catalina.core.ApplicationFilterChain.internalDoFilter(ApplicationFilterChain.java:197) at org.apache.catalina.core.ApplicationFilterChain.doFilter(ApplicationFilterChain.java:142) at coldfusion.monitor.event.MonitoringServletFilter.doFilter(MonitoringServletFilter.java:46) at coldfusion.bootstrap.BootstrapFilter.doFilter(BootstrapFilter.java:47) at org.apache.catalina.core.ApplicationFilterChain.internalDoFilter(ApplicationFilterChain.java:166) at org.apache.catalina.core.ApplicationFilterChain.doFilter(ApplicationFilterChain.java:142) at org.apache.catalina.core.StandardWrapperValve.invoke(StandardWrapperValve.java:166) at org.apache.catalina.core.StandardContextValve.invoke(StandardContextValve.java:88) at org.apache.catalina.authenticator.AuthenticatorBase.invoke(AuthenticatorBase.java:481) at org.apache.catalina.core.StandardHostValve.invoke(StandardHostValve.java:127) at org.apache.catalina.valves.ErrorReportValve.invoke(ErrorReportValve.java:83) at org.apache.catalina.core.StandardEngineValve.invoke(StandardEngineValve.java:72) at org.apache.catalina.connector.CoyoteAdapter.service(CoyoteAdapter.java:357) at org.apache.coyote.ajp.AjpProcessor.service(AjpProcessor.java:452) at org.apache.coyote.AbstractProcessorLight.process(AbstractProcessorLight.java:63) at org.apache.coyote.AbstractProtocol$ConnectionHandler.process(AbstractProtocol.java:935) at org.apache.tomcat.util.net.NioEndpoint$SocketProcessor.doRun(NioEndpoint.java:1831) at org.apache.tomcat.util.net.SocketProcessorBase.run(SocketProcessorBase.java:52) at org.apache.tomcat.util.threads.ThreadPoolExecutor.runWorker(ThreadPoolExecutor.java:973) at org.apache.tomcat.util.threads.ThreadPoolExecutor$Worker.run(ThreadPoolExecutor.java:491) at org.apache.tomcat.util.threads.TaskThread$WrappingRunnable.run(TaskThread.java:63) at java.base/java.lang.Thread.run(Thread.java:834) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
| Suppressed |
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| TagContext |
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| Type | Application | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
| msg | File or directory E:\inetpub\wwwroot\test_idtools_org\tools\1077\images\thumbnails\retinue_of_marked,_clipped_queen_R_Snyder_1.jpg does not exist. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
| Form - struct | |
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| TEXT1 | [empty string] |
| TEXT2 | [empty string] |
| TEXT3 | [empty string] |
| URL - struct | |
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| entityID | 59237 |
| packageID | 2268 |
| pageID | 1772 |
| All Arguments - struct | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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| EVENTNAME | [empty string] | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
| EXCEPTION |
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